Level 1
[Trait] Fever Pitch
Increases Frenzy's Attack Speed bonus to 10%.
[Q/W] Grooved Spines
Increases the range of Poisoned Spine and Spine Barrage by 2.
[E] Nutrient Absorption
While Burrowed, Apex regenerates 2% of his maximum Health every second.
? Quest: Collect Regeneration Globes to increase Apex's Health Regeneration while burrowed by 0.15% of his maximum Health per second, up to 3%.
? Reward: After collecting 20 Regeneration Globes, Apex always regenerates 2% of his maximum Health every second.
[Passive] Muscular Augments
Gain 10% increased Movement Speed.
Each stack of Frenzy grants 1% additional Movement Speed.
Level 4
[Trait/Basic Attack] Pressurized Ducts
While at 10 stacks of Frenzy, Apex's Attack Range is increased to 7.7.
? Quest: Make 80 Basic Attacks against enemy Heroes while at 10 stacks of Frenzy.
? Reward: Apex's Basic Attack range is permanently increased to 7.7.
[Q] Paralytic Barbs
Poisoned Spine Slows the target by 60%, decaying over 3 seconds.
[W] Serrated Spines
Spine Barrage deals 100% additional damage to non-Heroes.
[Passive] Blood Craze
Basic Attacks against enemy Heroes reduce Apex's Basic Ability cooldowns by 0.5 seconds.
Level 7
[Trait] Frenzied Endurance
Basic Attacks heal Apex for 2% of the damage they deal for each stack of Frenzy he has, increased to 4% per stack against Heroes.
[W] Spine Storm
Spine Barrage gains an additional charge and its charge cooldown is reduced to 0.5 seconds.
[Q] Parasitic Venom
Poisoned Spine's poison heals Apex for 25% of the damage it deals to enemies, increased to 75% against Heroes.
Level 13
[Trait] Hunter's Instinct
Every consecutive Basic Attack against the same enemy Hero within 4 seconds grants 1 additional stack of Frenzy per attack, up to 4 per attack.
[Q] Immune Suppressant
Increases Poisoned Spine's poison duration by 1 second.
Poisoned Spine and Basic Attacks against enemies poisoned by Poisoned Spine reduce the healing they receive by 30% for 2 seconds.
[W] Tangler Spores
Tentacles sprout from the ground where Spine Barrage lands after 1 second, dealing 60 damage per second and Slowing enemies by 40% for 3 seconds.
[E] Adaptive Talons
Reduces Burrow's channel period to 0.25 seconds.
Apex can now move at 60% Movement Speed while Burrowed, but disturbs the ground as he moves and makes his location visible.
Level 16
[Q] Aggressive Neurotoxin
Poisoned Spine's poison deals additional damage to Heroes equal to 0.5% of their maximum Health per second.
Poisoned Spine's poison deals additional damage to Heroes equal to 1% of their maximum Health each time it is refreshed or reapplied.
[W] Spinestorm
Casting Spine Barrage empowers Apex's next 2 Basic Attacks within 6 seconds. These attacks can stack across consecutive casts of Spine Barrage.
Apex's empowered attacks launches a mini-Spine Barrage at the target's location, dealing 50% of Spine Barrage's damage in a radius of 2 around the target.
[Passive] Razorspines
All of Apex's abilities and Basic Attacks ignore Armor.
Poisoned Spine and Basic Attacks now damage and pierce all enemies in their path out to their maximum range.
[Active] Landshark
Activate while Burrowed to tunnel quickly through the ground in a targeted direction, dealing 200 damage to enemies within 2 range of Apex's path.
Level 20
[Trait] Mass Frenzy
Half of Frenzy's Attack Speed bonus is shared with all allied Minions and non-Elite Monsters and Mercenaries within 8 range.
[Trait] Hunter-Killer
? Quest: Killing an enemy Hero grants 1 permanent stack of Frenzy, up to 10.
? Reward: After killing 10 enemy Heroes, Frenzy's cooldown is set to 60 seconds and it now reveals the location of all enemy Heroes on the map for 4 seconds.
[R1] Impaling Tentacle
At a targeted point along Seismic Spines' length a single tentacle erupts from the ground, dealing an additional 280 damage to enemies in a radius of 2.
[R2] Primal Rage
Taking fatal damage while Wild Mutation is active grants Unkillable for 3 seconds. This can only happen once per cast of Wild Mutation.
Killing an enemy Hero during Wild Mutation reduces its cooldown to 5 seconds.
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